I expect that the car would slow down as soon as he lifted his foot off the accelerator.
Just want to get at the braking claim for now, and the rest later.
"I wonder if the Z250s is where Greer inadvertently began to ease
his foot off the accelerator. That would begin the process that
caused the car to slow down by time it reached the Z290s."
Well, I didn't say (above) Greer completely lifted his foot off the accelerator. I speculated Greer may have slowly eased his foot off, beginning in the Z250s, when I believe both Kellerman and Greer looked back, possibly responding to the same causation coming from behind. Such a gradual lifting off the accelerator would gradually slow down the car.
This is from an article by Chuck Marler in the old newsletter "Fourth Decade" (May 1994):
"Based upon the government's measurements, the limousine only reduced
its speed 1.2 miles per hour between frames 225 through 255 and frames
255 through 313. When one views the Zapruder film without the close–up
enhancements, the limousine reduces its speed significantly between frames
255 and 313—substantially more than the 1.2 miles per hour. This obvious
reduction of speed prior to frame 313 also occurs as the limousine is
becoming more perpendicular to the location of Abraham Zapruder which
should visually appear to be going faster even if there wasn't a reduction
in speed."
However, there is evidence in the Muchmore film that the brake lights came on as Clint Hill ran to the limo:
In the above frame, the left brake light is not on.
The right brake light is on in the above two frames but in the following frame as Clint Hill reaches the limo the brake like is off.
Those aren't brake lights that are lit up. They're "wrap-around" tail-light lenses that allow ambient light to shine through.
Whether one tail-light or both (or none) were artificially bright depended on the viewer's position and amount and direction of ambient light. Now you'll argue Ford Lincoln wouldn't allow such a tail-light design because it would be a traffic hazard.
Tail-light in M19 and M45 (the frame you supplied in your post) are about equal in brightness. Neither appear to be lit by a bulb.
Physicist Luis Alvarez did an analysis of the limo speed ("A Physicist Examines the Kennedy Assassination Film", American Journal of Physics, Sept. 1976, Vol 44, No. 9, p. 813 at p. 825 and determined that it began to slow down around z300:
which corresponds to the time that Clint Hill was running as seen in Muchmore.
Looks like the slowing event Alvarez presented was centered on Z300. Alvarez wrote:
"The heavy car decelerated suddenly for about 0.5 sec (10 frames),
centered at about frame 299, reducing its speed from about 12 mph
to about 8 mph." (or frame 300 in his graph)
So now we look for a brake light being on at Z295. There is a still of Muchmore frame M24 (Z295) in Robin's Gallery. As it happens, it's one of the clearest frames.
In the animation, see if the limousine tail-lights come on before the tail-light passes the little boy. That's when braking must occur if Greer braked to account for the slowdown Alvarez centered on Z300.